| 1. |
- Paterson, John R., et al.
(författare)
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An early Cambrian faunule from the Koolywurtie Limestone Member (Parara Limestone), Yorke Peninsula, South Australia and its biostratigraphic significance
- 2007
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Ingår i: Memoirs of the Association of Australasian Palaeontologists. - 0810-8889. ; 34, s. 131-146
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Tidskriftsartikel (refereegranskat)abstract
- An early Cambrian shelly faunule from the Koolywurtie Limestone Member (Parara Limestone) at Teppers Knoll on Yorke Peninsula, South Australia, includes lingulate brachiopods, a variety of small shelly fossils such as tommotiids, hyolithelminth tubes and orthothecides, chancelloriid sclerites, sponge spicules, in addition to the oldest known occurrence of the redlichioid trilobite Xela. Correlation using faunas from other South Australian successions suggests that the member, in addition to the archaeocyathan fauna referred to as the ‘Syringocnema favus beds’, are early Botoman (pre-Pararaia janeae Zone) in age.
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| 2. |
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| 3. |
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| 4. |
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| 5. |
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| 6. |
- Balthasar, Uwe, et al.
(författare)
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Homologous skeletal secretion in tommotiids and brachiopods
- 2009
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Ingår i: Geology. - 0091-7613 .- 1943-2682. ; 37:12, s. 1143-1146
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Tidskriftsartikel (refereegranskat)abstract
- Tommotiids are distinctive components of the early Cambrian small shelly fauna, almost invariably represented by isolated phosphatic sclerites derived from a multielement protective cover (scleritome). The unusual range of tommotiid sclerite morphologies and unknown construction of the scleritome have severely hampered our understanding of their phylogenetic affinities. However, recent description of rare, articulated scleritome material belonging to the tommotiid genera Eccentrotheca and Paterimitra support the hypothesis that some tommotiids fall within the stem group of the lophophorate phyla Phoronida and Brachiopoda and that at least some tommotiid sclerites are homologous precursors of the shells of organophosphatic brachiopods. Here we show that the shell microstructure of Eccentrotheca and Paterimitra share substantial similarities with paterinid brachiopods. While paterinids possess an overall brachiopod morphology, their microstructure appears more similar to Eccentrotheca and Paterimitra than to nonpaterinate lingulids. These findings strongly support the existence of a brachiopod total group that is solidly rooted within tommotiids, and identify the organophosphatic skeletal composition as plesiomorphic with calcareous shells as derived. The microstructural changes of the proposed tommotiid-brachiopod transition probably reflect an adaptation to fluctuating food and phosphorous intake that came with the switch to a sessile life style at the base of the tommotiid clade.
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| 7. |
- Betts, Marissa, J., et al.
(författare)
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A new lower Cambrian shelly fossil biostratigraphy for South Australia
- 2016
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Ingår i: Gondwana Research. - : Elsevier BV. - 1342-937X .- 1878-0571. ; 36, s. 163-195
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Tidskriftsartikel (refereegranskat)abstract
- Definition of early Cambrian chronostratigraphic boundaries is problematic with many subdivisions stillawaiting ratification. Integrated multi-proxy data from well-resolved regional-scale schemes are ultimately the key to resolving broader issues of global correlationwithin the Cambrian. In Australia, early Cambrian biostratigraphy has been based predominantly on trilobites. Phosphatic shelly fauna have great potential as biostratigraphic tools, especially in pre-trilobitic strata because they are widespread and readily preserved, but they have remained underutilised. Here we demonstrate their value in a new biostratigraphic scheme for the early Cambrian of South Australia using a diverse shelly fauna including tommotiids, brachiopods, molluscs and bradoriids. Biostratigraphic data are derived from ten measured stratigraphic sections across the Arrowie Basin, targeting Hawker Group carbonates including the Wilkawillina, Wirrapowie and Ajax limestones and the Mernmerna Formation. The stratigraphic ranges of shelly fossils are predictable and repeatable across the Arrowie Basin, allowing three discrete shelly biozones to be identified, spanning Terreneuvian, Stage 2 to Series 2, Stages 3–4. The Kulparina rostrata Zone (new) and part of the overlyingMicrina etheridgei Zone (new) are pre-trilobitic (predominantly Terreneuvian). The Cambrian Series 2, Stage 3 Dailyatia odyssei Zone (new) features a very diverse shelly fauna and will be described in detail in a separate publication. These zones provide robust means to correlate Terreneuvian–Series 2 successions in neighbouring coeval basins in Australia, particularly the Stansbury Basin. Wider correlation is possible throughout East Gondwana, and especially with South China.
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| 8. |
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| 9. |
- Betts, Marissa, J., et al.
(författare)
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Early Cambrian chronostratigraphy and geochronology of South Australia
- 2018
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Ingår i: Earth-Science Reviews. - : Elsevier BV. - 0012-8252 .- 1872-6828. ; 185, s. 498-543
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Tidskriftsartikel (refereegranskat)abstract
- The most successful chronostratigraphic correlation methods enlist multiple proxies such as biostratigraphy and chemostratigraphy to constrain the timing of globally important bio- and geo-events. Here we present the first regional, high-resolution shelly fossil biostratigraphy integrated with δ13C chemostratigraphy (and corresponding δ18O data) from the traditional lower Cambrian (Terreneuvian and provisional Cambrian Series 2) of South Australia. The global ZHUCE, SHICE, positive excursions II and III and the CARE are captured in lower Cambrian successions from the Arrowie and Stansbury basins. The South Australian shelly fossil biostratigraphy has a consistent relationship with the δ13C results, bolstering interpretation, identification and correlation of the excursions. Positive excursion II straddles the boundary between the Kulparina rostrata and Micrina etheridgei zones, and the CARE straddles the boundary between the M. etheridgei and Dailyatia odyssei zones, peaking in the lower parts of the latter zone. New CA-TIMS zircon dates from the upper Hawker Group and Billy Creek Formation provide geochronologic calibration points for the upper D. odyssei Zone and corresponding chemostratigraphic curve, embedding the lower Cambrian successions from South Australia into a global chronostratigraphic context. This multi-proxy investigation demonstrates the power of integrated methods for developing regional biostratigraphic schemes and facilitating robust global correlation of lower Cambrian successions from South Australia (part of East Gondwana) with coeval terranes on other Cambrian palaeocontinents, including South and North China, Siberia, Laurentia, Avalonia and West Gondwana.
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| 10. |
- Betts, Marissa, J., et al.
(författare)
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Global correlation of the early Cambrian of South Australia: Shelly faunaof the Dailyatia odyssei Zone
- 2017
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Ingår i: Gondwana Research. - Amsterdam : Elsevier BV. - 1342-937X .- 1878-0571. ; 46, s. 240-279
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Tidskriftsartikel (refereegranskat)abstract
- A lack of well resolved biostratigraphic data has prevented robust regional and global correlation of lower Cambriansuccessions from South Australia. A new early Cambrian biostratigraphy, based on data derived from 21measured stratigraphic sections and drill cores (11 described herein) reveals the abundance and diversity ofshelly fauna from the Arrowie Basin, and the value of early Cambrian “small shelly fossils” (SSF) for biostratigraphicstudies. Here we examine shelly fauna associated with the youngest of three recently establishedbiozones, the Dailyatia odyssei Taxon Range Zone (hereafter D. odyssei Zone), and their correlative potential.The D. odyssei Zone features a diverse suite of tommotiids, organophosphatic brachiopods, bradoriid arthropods,molluscs and phosphatic problematica. This fauna permits strong correlation (often at species-level) with othermajor early Cambrian terranes, particularly Antarctica, South China and Laurentia, and suggest a Cambrian Series2, Stages 3–4 age for the D. odyssei Zone. Bradoriids have proven to be useful biostratigraphic tools. Four newspeciesand three new genera are described herein: Acutobalteus sinuosus gen. et sp. nov., Eozhexiella adnyamathanha gen. etsp. nov., Manawarra jonesi gen. et sp. nov. and Mongolitubulus descensus sp. nov. The description of Eohadrotreta sp.cf. zhenbaensis represents the first occurrence of the acrotretoid brachiopod Eohadrotreta from Australia.
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| 11. |
- Betts, Marissa J, et al.
(författare)
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Integrated chronostratigraphy of the lower Cambrian Byrd Group, Transantarctic Mountains
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Annan publikation (övrigt vetenskapligt/konstnärligt)abstract
- Antarctica and Australia were sutured together straddling the equator during the major pulse of animal biodiversification associated with the Cambrian radiation. However, lack of detailed systematic sampling of lower Cambrian sedimentary packages from Antarctica has significantly impeded precise age determination and correlation with other Cambrian paleocontinents, especially with other parts of East Gondwana. Here were present new, integrated biostratigraphic and chemostratigraphic (δ13C isotopes) data from three stratigraphic sections measured through autochthonous shallow water carbonates (including archaeocyath-microbial bioherms) from the lower Cambrian Byrd Group in the Transantarctic Mountains. Recovered shelly fossil assemblages (brachiopods, tommotiids, molluscs, trilobites) from the Holyoake and Churchill Ranges include conspecific taxa previously described from Hawker Group rocks in the Arrowie Basin of South Australia facilitating direct correlation with the upper Dailyatia odyssei biozone. Synchronous chemostratigraphic data capture a distinctive positive ?13C excursion in the Churchill Range interpreted as the global Mingxinsi Carbon Isotope Excursion (MICE) peak. A succeeding gradual negative ?13C excursion captured in the Churchill and Holyoake sections is interpreted as the global Archaeocyathid Extinction Carbon Isotope Excursion (AECE) event. There is no chemostratigraphic evidence for the large Redlichiid-Olenellid Extinction Carbon Isotope Excursion (ROECE) negative event that straddles and defines the Cambrian Stage 4 – Miaolingian boundary. Hence, the integrated faunal and new chemostratigraphic data presented herein strongly support a Cambrian Stage 4 age for the upper Shackleton Limestone – Holyoake Formation – Starshot Formation succession of the Byrd Group.
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| 12. |
- Betts, Marissa, J., et al.
(författare)
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Shelly fossils from the lower Cambrian White Point Conglomerate, Kangaroo Island, South Australia
- 2019
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Ingår i: Acta Palaeontologica Polonica. - : Polska Akademia Nauk Instytut Paleobiologii (Institute of Paleobiology, Polish Academy of Sciences). - 0567-7920 .- 1732-2421. ; 64:3, s. 489-522
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Tidskriftsartikel (refereegranskat)abstract
- The lower Cambrian (Series 2) White Point Conglomerate (WPC) on Kangaroo Island, South Australia contains exoticclasts representing a diverse array of lithologies, including metamorphics, chert, sandstone, and abundant carbonates,notably archaeocyath-rich bioclastic limestone. Acetic acid digestion of the WPC bioclastic limestone clasts reveals adiverse shelly fauna. This assemblage includes abundant organophosphatic brachiopods such as Cordatia erinae Brockand Claybourn gen. et sp. nov., Curdus pararaensis, Eodicellomus elkaniformiis, Eohadrotreta sp. cf. E. zhenbaensis,Eoobolus sp., Kyrshabaktella davidii, and Schizopholis yorkensis. Additional shelly taxa include the solenopleurid trilobiteTrachoparia? sp., the tommotiids Dailyatia odyssei, Dailyatia decobruta Betts sp. nov., Kelanella sp., and Lapworthellafasciculata, spines of the bradoriid arthropod Mongolitubulus squamifer, and several problematica, such as Stoibostrombuscrenulatus and a variety of tubular forms. The upper age limit for the WPC is constrained by biostratigraphic data fromthe overlying Marsden Sandstone and Emu Bay Shale, which are no younger than the Pararaia janeae Trilobite Zone(Cambrian Series 2, Stage 4). The shelly fossil assemblage from the WPC limestone clasts indicates an upper Dailyatiaodyssei Zone (= Pararaia tatei to lower P. janeae trilobite zones), equivalent to the Atdabanian–early Botoman of theSiberian scheme. This contrasts with the previously suggested late Botoman age for the limestone clasts, based on the diversearchaeocyath assemblage. The minor age difference between the WPC and its fossiliferous limestone clasts suggestsrelatively rapid reworking of biohermal buildups during tectonically-active phases of deposition in the Stansbury Basin.
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| 13. |
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| 14. |
- Claybourn, Thomas M., 1989-
(författare)
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Biostratigraphy and Systematics of Cambrian Small Shelly Fossils from East Antarctica and South Australia
- 2021
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Doktorsavhandling (övrigt vetenskapligt/konstnärligt)abstract
- The remote lower Cambrian Byrd group of sedimentary rocks from East Antarctica has been studied intermittently since its discovery over a century ago. Previous insights into the trilobites and archaeocyaths indicated a close correlation to the sedimentary sequences of South Australia. The lowest unit of the Byrd Group is the fossiliferous Shackleton Limestone which overlies the Neoproterozoic metamorphic rocks of the Beardmore Group and is representative of a long period of carbonate shelf formation on a passive margin with the palaeo-Pacific. This was truncated by marine transgression and the deposition of the deeper-water calcareous siltstones of the fossiliferous Holyoake Formation. This is overlain by the Starshot Formation and all three units are cross-cut by the Douglas Conglomerate, marking the start of a collisional tectonic regime between the East Gondwana and palaeo-Pacific plates. The first systematically sampled and analysed sections through the carbonate Shackleton Limestone and argillaceous Holyoake Formation has yielded a new fauna of small primarily phosphatic and secondarily phosphatised shelly fossils. The abundant molluscs, brachiopods and tommotiids are reported here. These findings are ideal for correlating this section to the fossil biozones of South Australia, including the Dailyatia odyssei small shelly fossil Zone and the Pararaia janeae trilobite Zone. Chemostratigraphic data from three sections preserve the profiles of two major stable carbon isotope excursions: the Mingxinsi Carbon Isotope Excursion and the Archaeocyathid Extinction Carbon isotope Excursion. The combination of these two lines of evidence are a strong indicator for Cambrian Series 2, early-mid Stage 4. This is corroborated by newly described D. odyssei-P. janeae Zone small shelly fossils from the carbonate clasts from the Cambrian Stage 4 White Point Conglomerate of South Australia which bear strong similarity to the fauna of the Shackleton Limestone. Palaeobiogeographically the fauna recovered from the Byrd Group is similar to the East Gondwanan region of South Australia, with similar brachiopod assemblages to those recovered from the Xinji Formation of North China and similar molluscan assemblages to the Bastion Formation of North-East Greenland.
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| 15. |
- Claybourn, Thomas M, et al.
(författare)
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Cambrian Series 2, Stage 3–4 micromolluscs from the Shackleton Limestone, Central Transantarctic Mountains, East Antarctica
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Annan publikation (övrigt vetenskapligt/konstnärligt)abstract
- An assemblage of lower Cambrian (Series 2, Stages 3–4) molluscan fossils from the Shackleton Limestone, Transantarctic Mountains, East Antarctica is formally described and illustrated. The assemblage is composed of one bivalve and ten helcionelloid molluscs, including the first discovery of the genus Xinjispira outside North China. The new fauna shows some similarity to previously described micromollusc assemblages from lower Cambrian glacial erratics from the Antarctic Peninsula. The fauna of the Shackleton Limestone is mainly composed of steinkerns, and of relatively low diversity, but the presence of key taxa, including the molluscs Mackinnonia rostrata and Pojetaia runnegari and the tommotiid Dailyatia odyssei strengthens correlation to the newly defined Dailyatia oddysei Zone of South Australia
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| 16. |
- Gubanov, Alexander P., et al.
(författare)
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Anabarella australis (Mollusca, Helcionelloida) from the Lower Cambrian of Greenland
- 2004
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Ingår i: Geobios. - : Elsevier BV. - 0016-6995 .- 1777-5728. ; 37:6, s. 719-724
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Tidskriftsartikel (refereegranskat)abstract
- The Early Cambrian helcionelloid mollusc Anabarella australis is described from North-East Greenland, representing the second occurrence of the species outside of Australia. Other Australian molluscs of this age are known from many localities including North China, Siberia, Altai, Transbaikalia, southern Kazakhstan, Mongolia, eastern Germany and Spain. These records, supported now by A. australis, demonstrate the close proximity of continents in the Early Cambrian.
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| 17. |
- Holmer, Lars E., et al.
(författare)
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A stem group brachiopod from the Lower Cambrian - support for a Micrina (Halkieriid) ancestry.
- 2002
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Ingår i: Palaeontology. - : Wiley. - 0031-0239 .- 1475-4983. ; 45:5, s. 875-882
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Tidskriftsartikel (refereegranskat)abstract
- The shell structure of the Lower Cambrian Mickwitzia, a bilaterally symmetrical bivalve hitherto doubtfully assigned to the Brachiopoda, confirms that the genus shares characters with linguliform brachiopods. The columnar lamination of its organophosphatic shell is homologous with that characterizing acrotretides. The shell, however, is also pervaded by striated apatitic tubes indistinguishable from those permeating the sclerites of the problematic organophosphatic, laminar–shelled Micrina which is close to Halkieria. No crown group brachiopods have such tubes that are presumed to have contained setae. The presence of both these features in the Mickwitzia shell suggests that the stock is a stem group brachiopod with a halkieriid ancestry.
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| 18. |
- Holmer, Lars E., et al.
(författare)
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First record of a bivalved larval shell in Early Cambrian tommotiids and its phylogenetic significance
- 2011
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Ingår i: Palaeontology. - : Wiley. - 0031-0239 .- 1475-4983. ; 54:2, s. 235-239
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Tidskriftsartikel (refereegranskat)abstract
- Brachiopods are marine Lophotrochozoa whose soft parts are enclosed in a bivalved shell. Although brachiopods are represented by a rich record from the Early Cambrian to the present, the origin of their bivalved body plan remains controversial. The Early Cambrian organophosphatic tommotiids Micrina and Paterimitra from Australia have been proposed as stem brachiopods. Here, we describe their earliest ontogeny, indicating that tommotiids possessed bivalved planktotrophic larvae. The curious combinations of characters in Micrina and Paterimitra indicate that they may belong to the stems of the Linguliformea and Rhynchonelliformea, respectively. The bivalved shell of adult living brachiopods may represent a plesiomorphic character retained from planktic tommotiid larvae; the crown group body plan of the Brachiopoda may have evolved through the paedomorphic retention of a bivalved larval state.
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| 19. |
- Holmer, Lars E, et al.
(författare)
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First record of canaliform shell structure from the Lower Cambrian paterinate brachiopod Askepasma from South Australia
- 2006
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Ingår i: Memoirs of the Association of Australasian Palaeontologists. ; 32:01, s. 1-5
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Tidskriftsartikel (refereegranskat)abstract
- The organophosphatic shell structure of the paterinate (Cryptotretidae) brachiopod Askepasmasp. is described, based on new material from the Lower Cambrian of South Australia. Thelamellose shell is penetrated by phosphatic canals, oriented subparallel to the shell lamination;the canals were open to the exterior and most likely contained organic setae that would haveemerged between lamellae. This type of setigerous canal system was previously only knownfrom the stem group brachiopods Mickwitzia muralensis and M. cf. occidens as well as theenigmatic sclerites of the tannuolinid Micrina, which has been interpreted as a more basal stemgroup brachiopod.The organophosphatic shell structure of the paterinate (Cryptotretidae) brachiopod Askepasmasp. is described, based on new material from the Lower Cambrian of South Australia. Thelamellose shell is penetrated by phosphatic canals, oriented subparallel to the shell lamination;the canals were open to the exterior and most likely contained organic setae that would haveemerged between lamellae. This type of setigerous canal system was previously only knownfrom the stem group brachiopods Mickwitzia muralensis and M. cf. occidens as well as theenigmatic sclerites of the tannuolinid Micrina, which has been interpreted as a more basal stemgroup brachiopod.
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| 20. |
- Holmer, Lars E., et al.
(författare)
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The enigmatic early cambrian salanygolina : a stem group of rhynchonelliform chileate brachiopods?
- 2009
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Ingår i: Palaeontology. - : Wiley. - 0031-0239 .- 1475-4983. ; 52:1, s. 1-10
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Tidskriftsartikel (refereegranskat)abstract
- New material of the enigmatic brachiopod Salanygolina obliqua Ushatinskaya from the Early Cambrian of Mongolia shows that it has a colleplax- a triangular plate - in the umbonal perforation, which is enlarged by resorption. This structure is otherwise only known from the equally enigmatic Palaeozoic orders Chileida and Dictyonellida (Rhynchonelliformea, Chileata). The colleplax in Salanygolina is here considered to be homologous with that of the chileates. Salanygolina is also provided with a ridge-like pseudodeltidium, which is another chileate feature. Other characters of Salanygolina, like the radial arrangement of adductor muscle scars and postero-medially placed internal oblique muscles are characteristic of chileates, but also found in the paterinates. In contrast, mixoperipheral dorsal valves with low rudimentary interareas are well known in paterinates, but not yet recorded from chileates. Thus, Salanygolina shows a mosaic combination of morphologic characters, known both from the paterinates and chileates, indicating that it may represent a stem group of the rhynchonelliform chileate brachiopods. The laminar phosphatic secondary shell of Salanygolina is composed of closely packed and nearly identical hexagonal prisms, oriented with their long axis normal to the laminae in a honeycomb pattern. The prism walls appear to have originally been composed of organic membranes and might represent precursors of the organic sheaths of calcite fibers that are typical of calcitic shells with a fibrous microstructure.
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| 21. |
- Holmer, Lars Erik, 1960-, et al.
(författare)
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The Origin of the Brachiopod Body Plan
- 2009
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Ingår i: Internationl Conference on the Cambrian Explosion. - 9780981288512 ; , s. 541-541
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Konferensbidrag (refereegranskat)
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| 22. |
- Hu, Shixue, et al.
(författare)
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Soft-part preservation in a linguliform brachiopod from the lower Cambrian Wulongqing Formation (Guanshan Fauna) of Yunnan, South China
- 2010
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Ingår i: Acta Palaeontologica Polonica. - : Polska Akademia Nauk Instytut Paleobiologii (Institute of Paleobiology, Polish Academy of Sciences). - 0567-7920 .- 1732-2421. ; 55:3, s. 495-505
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Tidskriftsartikel (refereegranskat)abstract
- Linguliform brachiopods were important components of early Cambrian benthic communities. However, exceptionally preserved soft parts in Cambrian linguliform brachiopods are extremely sparse, and the most important findings are from the early Cambrian Chengjiang Konservat Lagerstatte of Kunming, southern China. Here we describe the first record of preserved soft-part anatomy in a linguli form brachiopod from the early Cambrian Guanshan fauna (Wu longging Formation, Palaeolenus Zone); a unit which is considerably younger than the Chengjiang fauna. The well preserved soft anatomy include linguliform pedicles, marginal setae and, in a few cases, an intact lophophore imprint. The pedicle has pronounced surface annulations, with its proximal-most part enclosing the apex of the ventral pseudointerarea; the pedicle is up to 51 mm long, corresponding to more than 4 times the sagittal length of the shell, and 12% of the maximum valve width. In details of their preservation, these new fossils exhibit striking similarities with the linguliforms from the older Chengjiang fauna, and all specimens are preserved in a compressed state as flattened impressions. The new linguliform has an elongate oval to subtriangular shell and an elongate triangular ventral pseudointerarea; the pedicle emerged from an apical foramen through a poorly preserved internal pedicle tube. The new linguliform is most similar to the mostly organic-shelled siphonotretoid-like brachiopod Acanthotretella spinosa, recently described from the classic middle Cambrian Burgess Shale Konservat Lagerstatte, British Columbia, Canada. The new species Acanthotretella decaius sp. nov. is described; it differs from A. spinosa in having a slightly thicker pedicle, and a larger and more rigid, probably partly mineralised shell, indicating that the mostly organic shell of A. spinosa may represent a secondary reduction of shell mineralisation. However, the spine-like setae of the new species are unfortunately poorly preserved only at the margin of the shell, but the new species is referred tentatively to the Superfamily Siphonotretoidea. The occurrence of A. decaius in the Guanshan fauna is the first lower Cambrian (Series 2, early Stage 4) record of both Acanthotretella and siphonotretoids, and it represents the first description of a lophophore and digestive tract from the siphonotretoid lineage.
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| 23. |
- Jacquet, Sarah, M., et al.
(författare)
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Watsonella crosbyi from the lower Cambrian (Terreneuvian, Stage 2) Normanville Group in South Australia
- 2016
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Ingår i: Geological Magazine. - 0016-7568 .- 1469-5081. ; , s. 1-17
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Tidskriftsartikel (refereegranskat)abstract
- Correlation of lower Cambrian strata is often confounded by provincialism of key fauna. The widespread occurrence of themicromollusc Watsonella crosbyi Grabau, 1900 is therefore an important biostratigraphic signpost with potential for international correlation of lower Cambrian successions. Previous correlations of W. crosbyi from Australia (Normanville Group) suggested an Atdabanian- to Botoman-equivalent age. However, in the upper part of the Mount Terrible Formation, stratigraphic ranges of W. crosbyi and Aldanella sp. cf. golubevi overlap prior to the incoming of vertically burrowed ‘piperock’, which is indicative of an age no earlier than Cambrian Stage 2. The stratigraphic range of W. crosbyi in the Normanville Group, South Australia correlates with the ranges of the taxon in China, France, Mongolia and Siberia (though not Newfoundland). The new Australian data add further support for considering the first occurrence of W. crosbyi a good potential candidate for defining the base of Cambrian Stage 2. The stratigraphic range of W. crosbyi through the lower Cambrian Normanville Group has been determined based on collections from measured sections. Although rare, W. crosbyi is part of an assemblage of micromolluscs including Bemella sp., Parailsanella sp. cf. murenica and a sinistral form of Aldanella (A. sp. cf. A. golubevi). Other fauna present include Australohalkieria sp., Eremactis mawsoni, chancelloriids and Cupitheca sp.
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| 24. |
- Kouchinsky, Artem, et al.
(författare)
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Early-middle Cambrian stratigraphy and faunas from northern Siberia
- 2022
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Ingår i: Acta Palaeontologica Polonica. - : Institute of Paleobiology, Polish Academy of Sciences. - 0567-7920 .- 1732-2421. ; 67:2, s. 341-464
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Tidskriftsartikel (refereegranskat)abstract
- New assemblages of skeletal fossils chemically extracted from carbonates of the Cambrian Stage 2-Drumian Stage are reported from the lower reaches of the Lena River as well as from the Khorbusuonka, Malaya Kuonamka, and Bol'shaya Kuonamka rivers in northern part of the Siberian Platform. The fauna studied with scanning electron microscopy includes brachiopods, molluscs, hyoliths, halkieriids, chancelloriids, tommotiids, lobopodians, palaeoscolecidans, bradoriids, echinoderms, anabaritids, hyolithelminths, and sponges showing similarity to previously described fossil assemblages from Siberia, Laurentia, and Gondwana. The material includes emended descriptions of Halkieria proboscidea, Hadimopanella knappologica, Archaeopetasus typicus, and first descriptions of Hadimopanella foveata Kouchinsky sp. nov. and Archaeopetasus pachybasalis Kouchinsky sp. nov. Affinity of Archaeopetasus to chancelloriids is suggested. Finding of an in-place operculum in a planispiral shell of Michniakia minuta enables reinterpretation of this form as a hyolith, not a mollusc. The cambroclavids Cambroclavus sp. and Zhijinites clavus and the earliest echinoderms belonging to the Rhombifera and Ctenocystoidea are reported respectively from the lower Botoman stage and Botoman-Toyonian transitional beds, correlated with Cambrian Stage 4. Carbon isotopes are analysed from sections of the Chuskuna (upper Kessyusa Group), Erkeket, Kuonamka, Olenyok, Yunkyulyabit-Yuryakh, Tyuser and Sekten formations. A major part of the delta C-13 record is obtained from the Cambrian Stage 4-Drumian Stage strata which remain incompletely characterised by chemostratigraphy. The Lower Anomocarioides limbataeformis Carbon isotope Excursion (LACE) from the Drumian Stage of the Khorbusuonka River is introduced herein. New chemostratigraphic data are used for regional and global correlation and facilitate study of the evolutionary development of animals and faunas through the "Cambrian explosion".
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| 25. |
- Larsson, Cecilia M
(författare)
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Early Cambrian Problematic Lophotrochozoans and Dilemmas of Scleritome Reconstructions
- 2012
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Doktorsavhandling (övrigt vetenskapligt/konstnärligt)abstract
- The emergence and radiation of metazoan body plans around the Precambrian/Cambrian boundary, some 500-600 million years ago, seems to be concordant with the appearance and diversification of preservable hard parts. Several Precambrian soft-bodied, multicellular organisms most likely represent stem-group bilaterians, but their fossil record is rather sparse. In contrast, the Cambrian fossil record is comparably rich – comprising hard part, trace fossil and delicate soft tissue preservation – and most animal phyla that we know of today had evolved by the end of the Cambrian. Consequently, this time represents an important period in the early evolution of metazoan life forms. Most skeletal remnants of invertebrate organisms from this period are preserved in incomplete, disarticulated sclerite assemblages, and the true architecture of the original skeletal structure, the scleritome, may therefore be hard to discern. Many scleritomous taxa have been suggested to be members of the lophotrochozoan clade, while their exact position within this group remains unclear. Such taxa are often referred to as Problematica. This thesis deals with some problematic scleritomous early Cambrian lophotrochozoans, and as such also addresses the dilemmas of scleritome reconstructions. In the first part, completely disarticulated calcareous sclerites from the lower Cambrian of North Greenland are described as Trachyplax arctica. Hypothetical scleritome reconstruction alternatives and comparisons to other scleritome-bearing taxa are discussed, but the lack of articulated material obscures any satisfactory conclusions regarding phylogenetic affinities and the original morphology of the organism. The other part of the thesis focuses on some minute, organophosphatic scleritomous metazoans, tommotiids, found in lower Cambrian limestone successions in South Australia – Paterimitra pyramidalis and Kulparina rostrata – their scleritome architecture and their phylogenetic relationship with paterinid brachiopods. The oldest brachiopod from South Australia, Askepasma saproconcha, and the slightly younger Askepasma toddense are also described and discussed. Based on articulated specimens, recently described partial scleritomes of the tommotiid Eccentrotheca helenia and similarities in shell ultrastructure with both Eccentrotheca and Askepasma, Paterimitra is interpreted as a stem-group brachiopod and reconstructed as a bilaterally symmetrical, sessile, filter feeder with a tubular/conical scleritome. The morphological similarities with Paterimitra point in the same direction for the slightly older Kulparina.
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| 26. |
- Li, Luoyang, et al.
(författare)
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Fibrous or Prismatic? A Comparison of the Lamello-Fibrillar Nacre in Early Cambrian and Modern Lophotrochozoans
- 2023
-
Ingår i: Biology. - : MDPI. - 2079-7737. ; 12:1, s. 113-113
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Tidskriftsartikel (refereegranskat)abstract
- The Precambrian–Cambrian interval saw the first appearance of disparate modern metazoan phyla equipped with a wide array of mineralized exo- and endo-skeletons. However, the current knowledge of this remarkable metazoan skeletonization bio-event and its environmental interactions is limited because uncertainties have persisted in determining the mineralogy, microstructure, and hierarchical complexity of these earliest animal skeletons. This study characterizes in detail a previously poorly understood fibrous microstructure—the lamello-fibrillar (LF) nacre—in early Cambrian mollusk and hyolith shells and compares it with shell microstructures in modern counterparts (coleoid cuttlebones and serpulid tubes). This comparative study highlights key differences in the LF nacre amongst different lophotrochozoan groups in terms of mineralogical compositions and architectural organization of crystals. The results demonstrate that the LF nacre is a microstructural motif confined to the Mollusca. This study demonstrates that similar fibrous microstructure in Cambrian mollusks and hyoliths actually represent a primitive type of prismatic microstructure constituted of calcitic prisms. Revision of these fibrous microstructures in Cambrian fossils demonstrates that calcitic shells are prevalent in the so-called aragonite sea of the earliest Cambrian. This has important implications for understanding the relationship between seawater chemistry and skeletal mineralogy at the time when skeletons were first acquired by early lophotrochozoan biomineralizers.
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| 27. |
- Li, Luoyang, et al.
(författare)
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Revisiting the molluscan fauna from the Cambrian (Series 2, stages 3–4) Xinji Formation of North China
- 2021
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Ingår i: Papers in Palaeontology. - : John Wiley & Sons. - 2056-2799 .- 2056-2802. ; 7:1, s. 521-564
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Tidskriftsartikel (refereegranskat)abstract
- A diverse group of molluscs from the Cambrian Series 2, Stages 3–4 Xinji Formation of the North China Block (NCB) is described, based on more than 4500 specimens from three well-studied sections in Shaanxi and Henan provinces, along the southern and southwestern margin of the NCB. Twenty molluscan species are identified, including one bivalve, three stem group gastropods, and 16 additional helcionelloids. Among these, six helcionelloid species are reported from the NCB for the first time, and one new species, Parailsanella luonanensis sp. nov. is proposed. This diverse molluscan fauna shares a large number of species with contemporaneous faunas of South Australia (15), Antarctica (7), Laurentia (6), Siberia (3) and South China (1). Faunal similarities are even greater on a generic level. The striking similarities of the molluscan faunas of North China, South Australia and Antarctica strongly support the hypothesis that the NCB was situated close to Eastern Gondwana, most likely close to South Australia in the Cambrian Epoch 2. In addition, well-preserved shell attachment muscle scars were observed in the helcionelloid Figurina figurina, with two pairs of symmetrical, continuous, band-like muscle scars, which are obviously different from the musculature of both gastropods and monoplacophorans. Because of this unique musculature, these characteristic Cambrian cap-like molluscs are assigned to the Helcionelloida rather than to the Gastropoda or Monoplacophora. A diverse group of molluscs from the Cambrian Series 2, Stages 3–4 Xinji Formation of the North China Block (NCB) is described, based on more than 4500 specimens from three well-studied sections in Shaanxi and Henan provinces, along the southern and southwestern margin of the NCB. Twenty molluscan species are identified, including one bivalve, three stem group gastropods, and 16 additional helcionelloids. Among these, six helcionelloid species are reported from the NCB for the first time, and one new species, Parailsanella luonanensis sp. nov. is proposed. This diverse molluscan fauna shares a large number of species with contemporaneous faunas of South Australia (15), Antarctica (7), Laurentia (6), Siberia (3) and South China (1). Faunal similarities are even greater on a generic level. The striking similarities of the molluscan faunas of North China, South Australia and Antarctica strongly support the hypothesis that the NCB was situated close to Eastern Gondwana, most likely close to South Australia in the Cambrian Epoch 2. In addition, well-preserved shell attachment muscle scars were observed in the helcionelloid Figurina figurina, with two pairs of symmetrical, continuous, band-like muscle scars, which are obviously different from the musculature of both gastropods and monoplacophorans. Because of this unique musculature, these characteristic Cambrian cap-like molluscs are assigned to the Helcionelloida rather than to the Gastropoda or Monoplacophora.
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| 28. |
- Liu, Fan, et al.
(författare)
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A fresh look at the Hyolithid Doliutheca from the Early Cambrian (Stage 4) Shipai Formation of the Three Gorges Area, Hubei, South China
- 2022
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Ingår i: Biology. - : MDPI AG. - 2079-7737. ; 11:6, s. 875-875
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Tidskriftsartikel (refereegranskat)abstract
- New hyolith specimens from the early Cambrian (Stage 4) of the Three Gorges area, western Hubei Province are described and assigned to the species Doliutheca orientalis. Doliutheca are preserved in two taphonomic modes: casts in silty mudstone revealing gross morphology and some soft parts, and internal molds in calcareous pelites, which exhibit new morphological details of the conch and operculum. SEM and Micro-CT analyses show that Doliutheca preserve well-developed platy clavicles and cardinal processes on the interior of the operculum composed of rod-shaped tubular elements. This observation and the distinct cardinal and conical shields of the operculum indicate that Doliutheca could be placed within the Family Paramicrocornidae, most recently established as a group of hyoliths closely related to hyolithids.
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| 29. |
- Liu, Fan, et al.
(författare)
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Soft part preservation in hyolithids from the lower Cambrian (Stage 4) Guanshan Biota of South China and its implications
- 2021
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Ingår i: Palaeogeography, Palaeoclimatology, Palaeoecology. - : Elsevier BV. - 0031-0182 .- 1872-616X. ; 562, s. 110079-110079
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Tidskriftsartikel (refereegranskat)abstract
- Hyoliths, known globally from the Cambrian to the Permian are generally recognized as early lophotrochozoan animals of uncertain phylogenetic position. The calcareous exoskeleton of hyolithids, one of the two subgroups of hyoliths, consist of four elements including a conical conch, an external operculum, and a pair of curved enigmatic helens, but their ecology and internal morphology have long been considered controversial. Here we present for the first time, remarkably preserved hyolithids with helens and interior soft tissues from the Guanshan Biota (Cambrian, Stage 4) of Shijiangjun section, Wuding County in eastern Yunnan Province of South China. Preserved soft tissues and organs encompass the digestive tract, muscle scars and connective tissues between the operculum and conch. Three modes of soft part preservation in hyolithids from the Guanshan Biota are described: (1) preservation through pyritization, (2) sediment-infilling of guts, and (3) bacterial biofilm pseudomorphs (expressed as large internal hollow cavities), resulting from endogenous bacterial decay of soft parts. Such differential preservation of hyolith guts indicates different taphonomic modes for the soft parts of Cambrian animals in the Guanshan Biota. The newly collected material yields the oldest preserved guts in hyolithids from Konservat-biotas, and reveals new evidence for reconstructing hyolithid anatomy, such as the articulation mechanism of the opercula and the internal structures in the conch. Comparisons with younger wellpreserved hyolithid specimens indicate that the gut anatomy of hyolithids was evolutionarily conservative from the early Cambrian through to at least the Ordovician.
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| 30. |
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| 31. |
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| 32. |
- Paterson, John R., et al.
(författare)
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Oikozetetes from the early Cambrian of South Australia : implications for halkieriid affinities and functional morphology
- 2009
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Ingår i: Lethaia. - : Scandinavian University Press / Universitetsforlaget AS. - 0024-1164 .- 1502-3931. ; 42:2, s. 199-203
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Tidskriftsartikel (refereegranskat)abstract
- Paterson, J.R., Brock, G.A. & Skovsted, C.B. 2009: Oikozetetes from the early Cambrian of South Australia: implications for halkieriid affinities and functional morphology. Lethaia, Vol. 42, pp. 199-203 Shells of Oikozetetes and isolated halkieriid sclerites from a section of the lower Cambrian Mernmerna Formation in the Flinders Ranges, South Australia, are tentatively considered as being derived from the same scleritome. Details of shell morphology and the possible combination of biomineralized shell and sclerites suggest that Oikozetetes, if interpreted correctly, is closely related to Halkieria. A new interpretation of Oikozetetes shell morphology, in addition to the first report of paired muscle scars on the interior surface, sheds new light on the possible functional morphology of halkieriid shells and the means of attaching the shell to the body. The occurrence of Oikozetetes in South Australia extends its biostratigraphic range to the lower Cambrian and biogeographic range to East Gondwana.
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| 33. |
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| 34. |
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| 35. |
- Popov, Leonid E., et al.
(författare)
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Earliest ontogeny of Early Palaeozoic Craniiformea : implications for brachiopod phylogeny
- 2010
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Ingår i: Lethaia. - : Scandinavian University Press / Universitetsforlaget AS. - 0024-1164 .- 1502-3931. ; 43:3, s. 323-333
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Tidskriftsartikel (refereegranskat)abstract
- Well preserved specimens of the Early Palaeozoic craniiform brachiopods Orthisocrania and Craniops retain clear evidence of a lecithotrophic larval stage, indicating the loss of planktotrophy early in their phylogeny. The size of the earliest mineralized dorsal shell was <100 mu m across, and the well preserved shell structure in these fossil craniiforms allows their earliest ontogeny to be compared directly with that of living Novocrania, in which the first mineralized dorsal shell (metamorphic shell) is secreted only after settlement of the lecithotrophic larvae. Immediately outside this earliest shell (early post-metamorphic or brephic shell) and in the rest of the dorsal valve the primary layer in both fossil and living craniiforms has characteristic radially arranged laths, which are invariably lacking in the earliest dorsal shell. The ventral valve of the fossil specimens commonly preserves traces of an early attachment scar (cicatrix), which is equal in size to the dorsal metamorphic shell, and the brephic post-metamorphic ventral valve also has a primary shell with radially arranged laths. However, a primary shell with radial laths is completely lacking in the ventral valve of living Novocrania, indicating that heterochrony may have been involved in the origin of the encrusting mode of life in living craniids; the entire ventral valve of Recent craniids (with the possible exception of Neoancistrocrania) may correspond to the earliest attachment scar of some fossil taxa such as Orthisocrania. It is also probable that the unique absence of an inner mantle lobe as well as the absence of lobate cells in Novocrania could be the result of heterochronic changes. The dorsal valve of both fossil and living craniiforms has a marked outer growth ring, around 500 mu m across, marking the transition to the adult, and a significant change in regime of shell secretion. The earliest craniiform attachment is considered to be homologous to the unique attachment structures described recently in polytoechioids (e.g. Antigonambonites) and other members of the strophomenate clade. However, unlike the craniiforms, polytoechioids and strophomenates all have planktotrophic larvae, and planktotrophy is most probably a plesiomorphic character for all Brachiopoda.
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| 36. |
- Skovsted, Christian B., et al.
(författare)
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Early Cambrian brachiopods and other shelly fossils from the basal Kinzers Formation of Pennsylvania
- 2010
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Ingår i: Journal of Paleontology. - 0022-3360 .- 1937-2337. ; 84:4, s. 754-762
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Tidskriftsartikel (refereegranskat)abstract
- An assemblage of seventeen species of Small Shelly Fossils, dominated by the brachiopod Eothele tubulus and species of the mollusk Yochelcionella, is described from the basal Kinzers Formation of Thomasville, Pennsylvania. The occurrence extends southwards the distribution of an Early Cambrian fauna (Cambrian Series 2, Stage 4) that is otherwise characteristic of the eastern shelf of Laurentia from New York to Greenland. The poorly known acrothelid brachiopod Eothele tubulus is redescribed based on large collections of ventral valves. The shell structure of E. tubulus is characterized by orthogonal baculae, and represents the oldest known example of a baculate shell structure, indicating that this type of shell structure evolved already in the Early Cambrian.
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| 37. |
- Skovsted, Christian B., et al.
(författare)
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Early Cambrian brachiopods from North-East Greenland
- 2005
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Ingår i: Palaeontology. - : Wiley. - 0031-0239 .- 1475-4983. ; 48:2, s. 325-345
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Tidskriftsartikel (refereegranskat)abstract
- A diverse assemblage of late Early Cambrian brachiopods is described from the Bastion and Ella Island formations of North-East Greenland. The fauna includes nine species, representing all three extant brachiopod subphyla in addition to the stem group brachiopod Mickwitzia cf. occidens. Four linguliforms: Eoobolus priscus, Botsfordia caelata, Micromitra bella, Vandalotreta sp., three rynchonelliforms: Obolella crassa, Kutorgina reticulata, and an unidentified chileid plus a possible craniiform species occur. The fauna shows similarities to late Early Cambrian (Dyeran Stage) brachiopod faunas of eastern Canada and the United States, but also to faunas from the late Early Cambrian (Botomian–Toyonian equivalent) of Australia, Antarctica and Siberia
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| 38. |
- Skovsted, Christian B., et al.
(författare)
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Early Cambrian record of failed durophagy and shell repair in an epibenthic mollusc
- 2007
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Ingår i: Biology Letters. - : The Royal Society. - 1744-9561 .- 1744-957X. ; 3:3, s. 314-317
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Tidskriftsartikel (refereegranskat)abstract
- Predation is arguably one of the main driving forces of early metazoan evolution, yet the fossil record of predation during the Ediacaran–Early Cambrian transition is relatively poor. Here, we present direct evidence of failed durophagous (shell-breaking) predation and subsequent shell repair in the Early Cambrian (Botoman) epibenthic mollusc Marocella from the Mernmerna Formation and Oraparinna Shale in the Flinders Ranges, South Australia. This record pushes back the first appearance of durophagy on molluscs by approximately 40 Myr.
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| 39. |
- Skovsted, Christian B., et al.
(författare)
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First occurrence of a new Ocruranus-like helcionelloid mollusc from the lower Cambrian of East Gondwana
- 2012
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Ingår i: Gondwana Research. - : Elsevier BV. - 1342-937X .- 1878-0571. ; 22:1, s. 256-261
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Tidskriftsartikel (refereegranskat)abstract
- A new cap-shaped mollusc, Emargimantus angulatus gen. et sp. nov. is described from the Arrowie Basin of South Australia. The new species is closely comparable to mollusc species from South China and North-East Greenland previously described under the generic name Ocruranus Liu, a genus recently reinterpreted as a multiplated, possibly polyplacophoran mollusc. Emargimantus is interpreted as a univalved helcionelloid mollusc and differs from Ocruranus in both morphology and function. E. angulatus represents the first discovery of Ocruranus-like helcionelloids in the lower Cambrian of eastern Gondwana and demonstrates that these molluscs had a global distribution during the early Cambrian.
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| 40. |
- Skovsted, Christian B., et al.
(författare)
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Hyolithellus in life position from the Lower Cambrian of North Greenland
- 2011
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Ingår i: Journal of Paleontology. - : Cambridge University Press (CUP). - 0022-3360 .- 1937-2337. ; 85:1, s. 37-47
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Tidskriftsartikel (refereegranskat)abstract
- Tubular specimens belonging to Hyolithellus from silty dolostones of the basal Aftenstjernesø Formation of North Greenland may represent the first occurrence of this widespread Cambrian fossil in life position. A high proportion of preserved specimens are oriented normal to bedding with the tapering end of the tube down. Occasional undulations in the growth of the tubes indicate that the animal actively adjusted its growth to achieve a vertical orientation in relation to the sediment surface. Increasing thickness of the tube wall towards the tapering end shifted the center of mass downwards and resulted in greater stability in the sediment. The tube remained open at both ends throughout ontogeny; it was most likely secreted by an annelid-grade animal which pumped water into the sediment through the tube. Hyolithellus and similar tubular fossils from the Lower Cambrian probably represent stem group annelids.
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| 41. |
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| 42. |
- Skovsted, Christian B., et al.
(författare)
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Sclerite fusion in the problematic early Cambrian spine-like fossil Stoibostrombus from South Australia
- 2011
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Ingår i: Bulletin of Geosciences. - : Czech Geological Survey. - 1214-1119 .- 1802-8225. ; 86:3, s. 651-658
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Tidskriftsartikel (refereegranskat)abstract
- New collections of the problematic spine like fossil Stoibostrombus crenulatus Conway Morris & Bengtson from the Mt. Scott Range and Wilkawillina Gorge in the central Flinders Ranges, South Australia contain fused sclerite composites. In each fused specimen the spines are merged along their lateral margins and the orientation of the spines is almost identical. These new specimens confirm that Stoibostrombus spines were dermal sclerites, arranged in lateral pairs or transverse rows. The nature of the animal secreting the spines remains elusive, but available evidence suggest that it was an ecdysozoan animal, possibly a palaeoscolecid worm.
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| 43. |
- Skovsted, Christian B., et al.
(författare)
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Scleritome construction, biofacies, biostratigraphy and systematics of the tommotiid Eccentrotheca helenia sp. nov. from the Early Cambrian of South Australia
- 2011
-
Ingår i: Palaeontology. - : Wiley. - 0031-0239 .- 1475-4983. ; 54, s. 253-286
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Tidskriftsartikel (refereegranskat)abstract
- Large collections of Eccentrotheca helenia sp. nov. from the lower Cambrian Wilkawillina and Ajax limestones in the Arrowie Basin, South Australia, contain abundant low, cap-shaped and high, laterally compressed isolated sclerites in addition to partially articulated tubular specimens. The scleritome of Eccentrotheca helenia sp. nov. is fully described for the first time and shown to be formed by ontogenetic fusion of sclerites into successively stacked sclerite rings, forming a larger, tubular structure. The apical termination of the tube is highly variable, but is primarily constructed by low, cap-shaped sclerites and characterised by a central aperture of variable inclination. The adapical portion of the tube is predominantly constructed by high, laterally compressed sclerites, but individual sclerite rings can contain both cap-shaped and laterally compressed sclerites along with sclerites of intermediate morphology. The apical aperture presumably housed organic structures for attachment to a hard substrate, but the scleritome also occasionally preserves small lateral perforations between fused sclerites, which may have served to stabilise the scleritome by providing additional points of anchorage. In the Arrowie Basin, E. helenia is found in association with archaeocyath-microbial-spongiomorph-dominated bioherms and most likely inhabited pendant or cryptic habitats within these bioherms. Eccentrotheca-like sclerites form an integral part of the scleritomes of many tommotiids which may confuse taxonomic analysis. Sclerites previously assigned to 'E.' guano, consistently occur together with sclerites of Kulparina rostrata in stratigraphic intervals consistently older than strata hosting E. helenia. Rare fused specimens indicate that the sclerites of K. rostrata and 'E.' guano belong to the same scleritome.
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| 44. |
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| 45. |
- Skovsted, Christian B., et al.
(författare)
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Small shelly fossils from the argillaceous facies of the Lower Cambrian Forteau Formation of western Newfoundland
- 2007
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Ingår i: Acta Palaeontologica Polonica. - 0567-7920 .- 1732-2421. ; 52:4, s. 729-748
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Tidskriftsartikel (refereegranskat)abstract
- A diverse fauna of helcionelloid molluscs, hyoliths, and other small shelly fossils is described from limestone layers within the Forteau Formation of the Bonne Bay region in western Newfoundland. The fauna is dominated by internal moulds of various molluscs and tubular problematica, but also includes hyolith opercula, echinoderm ossicles, and other calcareous small shelly fossils preserved by phosphatisation. Originally organophosphatic shells are comparatively rare, but are represented by brachiopods, hyolithelminths, and tommotiids. The fauna is similar to other late Early Cambrian faunas from slope and outer shelf settings along the eastern margin of Laurentia and may be of middle Dyeran age. The similarity of these faunas indicates that at least by the late Early Cambrian, a distinctive and laterally continuous outer shelf fauna had evolved. The Forteau Formation also shares elements with faunas from other Early Cambrian provinces, strengthening ties between Laurentia and Australia, China, and Europe during the late Early Cambrian. Two new taxa of problematic fossil organisms are described, the conical Clavitella curvata gen. et sp. nov. and the wedge-shaped Sphenopteron boomerang gen. et sp. nov.
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| 46. |
- Skovsted, Christian B., et al.
(författare)
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The Early Cambrian (Botomian) stem group brachiopod Mickwitzia from Northeast Greenland
- 2003
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Ingår i: Acta Palaeontologica Polonica. - 0567-7920 .- 1732-2421. ; 48:1, s. 1-20
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Tidskriftsartikel (refereegranskat)abstract
- The problematic brachiopod Mickwitzia Schmidt, 1888 is re-described based on new material of M. cf. occidens Walcott, 1908 from the Early Cambrian (Botomian) Bastion and Ella Island formations of Northeast Greenland. Etched material demonstrates that Mickwitzia has a lingulid-like juvenile ("larval") shell with trails of nick-points, reflecting the movement of marginal setae. Juvenile and early mature ventral valves have a lingulid-like pseudointerarea with a pedicle groove. The shell of M. cf. occidens is only partially phosphatic, in particular around the juvenile-early mature shell in both valves. The phosphatic shell includes at least two types of cylindrical structures: (1) slender columns identical with the columns of acrotretoid brachiopods and (2) relatively thicker tubes which may be open to the exterior surface and have internal striations (on the ventral pseudointerarea). The striations are most likely imprints of microvilli and these tubes can be inferred to have contained setae. The thinner linguliform columns and thicker setigerous striated tubes are considered to be homologous with identical structures in the sellate and mitral sclerites of the problematic Micrina, which has been identified as a probable primitive stem group of the Brachiopoda. Mickwitzia represents a more derived member of the stem group Brachiopoda.
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| 47. |
- Skovsted, Christian B., 1974-
(författare)
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The Early Cambrian Fauna of North-East Greenland
- 2003
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Doktorsavhandling (övrigt vetenskapligt/konstnärligt)abstract
- Small shelly fossils are common in sediments of Early Cambrian age and include the earliest common representatives of metazoan animals with mineralized hard parts. The group includes fossils of very different morphology, composition and ultrastructure. They seem to represent skeletal remains of numerous animal groups, the biological affinities of which are largely unresolved. However, the wide geographic range of many forms has the potential to enhance biostratigraphic and palaeogeographic resolution in the Early Cambrian. The late Early Cambrian sequence of North-East Greenland has yielded an assemblage of more than 88 species of small shelly fossils, brachiopods and trilobites, indicative of a middle Dyeran age (Botoman equivalent). The recovered fossils include a number of species that are known from other Early Cambrian palaeocontinents, and particularly strong ties to late Early Cambrian faunas of Australia are documented. The many cosmopolitan taxa thus identified suggests a close juxtaposition of palaeocontinents at this time. The systematic affinity of many of these small shelly fossils is poorly understood, partly because of their fragmentary nature and poor preservation. However, new data from North-East Greenland improves our understanding of the function and biological affinity of certain taxa. Collections of the problematic fossil Mongolitubulus from North and North-East Greenland exhibit characters indicative of a defensive function as spines of bivalved arthropods, while species of the problematic genus Triplicatella represent the opercula of an unknown tubular shell, probably related to orthothecid hyoliths. The bivalved fossil Mickwitzia from North-East Greenland combines characters of linguliform brachiopods and sclerites of Micrina, a non-bivalved problematic form (halkieriid) from Australia. The combination suggests that Mickwitzia is a stem group brachiopod and strengthens arguments for a halkieriid ancestry of the brachiopod phylum.
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| 48. |
- Skovsted, Christian B, et al.
(författare)
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The Lower Cambrian brachiopod Kyrshabaktella and associated shelly fossils from the Harkless Formation, southern Nevada
- 2006
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Ingår i: GFF. - 1103-5897 .- 2000-0863. ; 128, s. 327-337
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Tidskriftsartikel (refereegranskat)abstract
- A fauna of brachiopods and small shelly fossils from the Lower Cambrian Harkless Formation of Esmeraldina County in southern Nevada is described. The brachiopod Kyrshabaktella sp. provides the first unequivocal record of lingulids in the Lower Cambrian of the Great Basin. The shells of Kyrshabaktella sp. are fragmentary, but otherwise well preserved and an acrotretid-like columnar shell structure is demonstrated. The phosphatic tubular fossil Hyolithellus insolitus is described from North America for the first time and Sphenothallus sp. provides the oldest record of this problematic genus in Laurentian strata. Other faunal elements include two additional linguliform brachiopods, echinoderm ossicles and sponge spicules.
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| 49. |
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| 50. |
- Skovsted, Christian B., et al.
(författare)
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The problematic early Cambrian fossil Tumulduria incomperta represents the detached ventral interarea of a paterinid brachiopod.
- 2014
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Ingår i: Acta Palaeontologica Polonica. - : Polska Akademia Nauk Instytut Paleobiologii (Institute of Paleobiology, Polish Academy of Sciences). - 0567-7920 .- 1732-2421. ; 59:2, s. 359-365
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Tidskriftsartikel (refereegranskat)abstract
- The organophosphatic early Cambrian (Terreneuvian, Cambrian Stage 2) fossil Tumulduria incomperta has been problematic ever since its original description in 1969. Comparison of abundant specimens from the Lower Cambrian of Siberia with co-occurring brachiopod valves show that T. incomperta represents the central portion of the ventral interarea of a paterinid brachiopod similar to Cryptotreta neguertchenensis, and that the domed central portion of typical Tumulduria specimens represents the ridge-like pseudodeltidium of the interarea.
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