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1.	Felde, Vivian A., et al. (författare) Compositional turnover and variation in Eemian pollen sequences in Europe 2020 Ingår i: Vegetation History and Archaeobotany. - : Springer Science and Business Media LLC. - 0939-6314 .- 1617-6278. ; 29:1, s. 101-109 Tidskriftsartikel (refereegranskat)abstract The Eemian interglacial represents a natural experiment on how past vegetation with negligible human impact responded to amplified temperature changes compared to the Holocene. Here, we assemble 47 carefully selected Eemian pollen sequences from Europe to explore geographical patterns of (1) total compositional turnover and total variation for each sequence and (2) stratigraphical turnover between samples within each sequence using detrended canonical correspondence analysis, multivariate regression trees, and principal curves. Our synthesis shows that turnover and variation are highest in central Europe (47-55 degrees N), low in southern Europe (south of 45 degrees N), and lowest in the north (above 60 degrees N). These results provide a basis for developing hypotheses about causes of vegetation change during the Eemian and their possible drivers.
2.	Van Meerbeeck, C. J., et al. (författare) The nature of MIS 3 stadial-interstadial transitions in Europe : New insights from model-data comparisons 2011 Ingår i: Quaternary Science Reviews. - : Elsevier BV. - 0277-3791 .- 1873-457X. ; 30:25-26, s. 3618-3637 Tidskriftsartikel (refereegranskat)abstract 15 abrupt warming transitions perturbed glacial climate in Greenland during Marine Isotope Stage 3 (MIS 3, 60-27 ka BP). One hypothesis states that the 8-16 degrees C warming between Greenland Stadials (GS) and Interstadials (GI) was caused by enhanced heat transport to the North Atlantic region after a resumption of the Atlantic Meridional Overturning Circulation (AMOC) from a weak or shutdown stadial mode. This hypothesis also predicts warming over Europe, a prediction poorly constrained by data due to the paucity of well-dated quantitative temperature records. We therefore use a new evidence from biotic proxies and a climate model simulation to study the characteristics of a GS-GI transition in continental Europe and the link to enhanced AMOC strength. We compare reconstructed climatic and vegetation changes between a stadial and subsequent interstadial - correlated to GS15 and GI14 (similar to 55 ka BP) - with a simulated AMOC resumption using a three-dimensional earth system model setup with early-MIS 3 boundary conditions. Over western Europe (12 degrees W-15 degrees E), we simulate twice the annual precipitation, a 17 degrees C warmer coldest month, a 8 degrees C warmer warmest month, 1300 degrees C-day more growing degree days with baseline 5 degrees C (GDD5) and potential vegetation allowing tree cover after the transition. However, the combined effect of frequent killing frosts, <20 mm summer precipitation and too few GDD5 after the transition suggest a northern tree limit lying at similar to 50 degrees N during GI14. With these 3 climatic limiting factors we provide a possible explanation for the absence of forests north of 48 degrees N during MIS 3 interstadials with mild summers. Finally, apart from a large model bias in warmest month surface air temperatures, our simulation is in reasonable agreement with reconstructed climatic and vegetation changes in Europe, thus further supporting the hypothesis.
3.	Alexanderson, Helena, et al. (författare) Optical dating of a Late Quaternary sediment sequence from Sokli, northern Finland 2008 Ingår i: Geochronometria. - : Walter de Gruyter GmbH. ; 32, s. 51-59 Tidskriftsartikel (refereegranskat)abstract Interstadial and non-glacial stadial sediments collected in boreholes from Sokli in northeastern Finland have been dated by optically stimulated luminescence on quartz and feldspar grains. The quartz OSL ages follow stratigraphic order, with one exception, and support the litho- and biostratigraphical correlation with the NW European mainland climate-stratigraphy and the marine oxygen-isotope stages. Feldspar IRSL dates generally overestimate the age, probably due to incomplete bleaching. The data show that during the last glacial cycle northeastern Finland was not glaciated until MIS 5b, around 90 ka. Interstadial conditions occurred around ~94 ka (MIS 5c), ~74-80 ka (MIS 5a) and 42-54 ka (MIS 3). The OSL ages have large standard errors mainly due to small sample sizes, relatively poor luminescence characteristics and uncertainties in dose-rate determinations.
4.	Bos, Johanna A. A., et al. (författare) Flora, vegetation and climate at Sokli, northeastern Fennoscandia, during the Weichselian Middle Pleniglacial 2009 Ingår i: Boreas. - : Wiley. - 0300-9483 .- 1502-3885. ; 38:2, s. 335-348 Tidskriftsartikel (refereegranskat)abstract A 2 m thick laminated lacustrine deposit of silt and clay recovered from the high-latitudinal site at Sokli (northern Finland) provides a unique mid-Weichselian fossil record for Fennoscandia. High-resolution botanical and zoological analyses of the lacustrine deposit allow detailed reconstruction of the regional vegetational development and of the history of the lake and the wetland ecosystem within the Sokli basin during the early part of the Weichselian Middle Pleniglacial (=equivalent to Marine Isotope Stage (MIS) 3). The inferred terrestrial vegetation represented by the Sokli MIS 3 sequence (so-called Tulppio Interstadial) was probably low-arctic tundra, treeless but with shrub elements including juniper, willow, dwarf birch, ericoids, lycopods and a rich herb flora with a variety of arctic-alpine taxa and heliophilous, pioneer elements. The presence of herbs such as Rubus chamaemorus, Epilobium palustre, Potentilla palustris and Sphagnum, Drepanocladus and other mosses suggests that the lake was fringed by wet meadows and peatlands or peaty telmatic communities. The distributional ranges of pine and tree birch were probably only a few hundred kilometres south or southeast of Sokli. This is concordant with evidence for the presence of boreal tree taxa during the MIS 3 in the Baltic countries and further east in Europe, but contradicts with the commonly inferred treeless tundra or grass-dominated steppe conditions in central Europe.
5.	Engels, Stefan, et al. (författare) A comparison of three Eurasian chironomid-climate calibration datasets on a W-E continentality gradient and the implications for quantitative temperature reconstructions 2014 Ingår i: Journal of Paleolimnology. - : Springer Science and Business Media LLC. - 0921-2728 .- 1573-0417. ; 51:4, s. 529-547 Tidskriftsartikel (refereegranskat)abstract Multiple regional chironomid-climate calibration datasets are available to reconstruct quantitatively July air temperatures from fossil chironomid assemblages. We examined the relationship between July air temperature and the 40 most common chironomid taxa in three independent Eurasian calibration (training) sets. The estimated temperature optimum of each chironomid taxon is systematically lower (by similar to 1-2 A degrees C) in a Norwegian calibration set compared to Finnish and Russian calibration sets. This result might partly be explained by the fact that the Norwegian calibration set extends further at the cold end of the temperature gradient. A difference in continentality between the Russian sites and the European sites might also contribute to this pattern. The number of taxa that show a statistically significant unimodal response to temperature is higher in the Norwegian calibration set (34 out of 40 taxa) compared to the modern Finnish (11 of 37 taxa; 3 common taxa absent) and the Russian calibration set (20 of 40 taxa), probably due to the longer temperature gradient incorporated in the Norwegian calibration set. We applied all three calibration sets to fossil chironomid assemblages from the high-latitude study site of Sokli (northeast Finland), a site with a unique series of lacustrine deposits covering (amongst others) the Holocene, part of early MIS 3 (at similar to 53 ka) and MIS 5d-c (at similar to 110-95 ka) and with independent proxy-records for comparison. In the early Holocene and during MIS 5c, the chironomid-based temperature inferences from all three inference models had similar values. Temperature reconstructions based on the Norwegian calibration set are 2-4 A degrees C lower for the late Holocene, early MIS 3 and MIS 5d than the inferred temperatures based on the other calibration sets. Although the lakes included in the Finnish calibration set are located closest to the site of Sokli, evaluation tests and a comparison with independent proxy data suggests that the Norwegian calibration set provides the most suitable analogues for reconstruction purposes for most of the fossil assemblages. Our results imply that when choosing a calibration set for quantitative climate reconstructions on glacial timescales, regional proximity of the fossil site may not be a sufficient basis, and the length of the temperature gradient of the calibration dataset and factors such as the continentality gradient covered by the calibration set must also be considered.
6.	Engels, Stefan, et al. (författare) Early Weichselian (MIS 5d and 5c) temperatures and environmental changes in northern Fennoscandia as recorded by chironomids and macroremains at Sokli, northeast Finland 2010 Ingår i: Boreas. - : Wiley. - 0300-9483 .- 1502-3885. ; 39:4, s. 689-704 Tidskriftsartikel (refereegranskat)abstract A 25-m-long sediment record spanning the time from the Eemian to the Holocene was recovered from Sokli, northeast Finland. This study focuses on a 6-m-long sediment interval that is dated to the Early Weichselian period (MIS 5d and 5c) and consists of lacustrine and fluvial deposits. Using chironomid remains, botanical and zoological macroremains as well as sediment lithology, we were able to reconstruct past changes in the environment, including climate. The results indicate that the site was situated on a flood-plain during the latter stages of MIS 5d (Herning Stadial) and that summer temperatures might have been similar to 6 degrees C lower than at present. Although this value should be treated with caution, as numerical analysis shows that it has a very poor fit-to-temperature, this low reconstructed value concurs with several other reconstructions that are available from western Europe. During MIS 5c (Brorup interstadial), the depositional environment changed into a lake system, initially with stratification of the water and subsequently with complete mixing and a strong influence of streams. Both chironomid-based and macroremain-based temperature inferences indicate past July air temperatures that were significantly higher than at present. This result is in contrast to other (low-resolution) reconstructions from northern Fennoscandia that indicate past temperatures 6-7 degrees C lower than present using fossil coleopteran assemblages. However, several central European sites indicate that there was a phase during the Brorup interstadial that was characterized by high summer temperatures, and a comparison between the high-resolution reconstructions from western Europe and the results presented in this study suggests that the north south July air temperature gradient between the mid- and high-latitudes was much weaker during the Brorup interstadial than it is at present. High solar insolation values (particularly the obliquity) during the Brorup interstadial might explain the low summer temperature gradient over the European continent. A return to fluvial conditions occurred in the upper parts of the sediment sequence, and, after a brief interval of gyttja deposition under cooling conditions, the site became glaciated during MIS 5b.
7.	Finné, Martin, et al. (författare) Last Interglacial Climate in Northern Sweden-Insights from a Speleothem Record 2019 Ingår i: Quaternary. - : MDPI AG. - 2571-550X. ; 2:3 Tidskriftsartikel (refereegranskat)abstract Continental records with absolute dates of the timing and progression of climatic conditions during the Last Interglacial (LIG) from northern Europe are rare. Speleothems from northern Europe have a large potential as archives for LIG environmental conditions since they were formed in sheltered environments and may be preserved beneath ice sheets. Here, we present delta C-13 and delta O-18 values from speleothem Kf-21, from Korallgrottan in Jamtland (northwest Sweden). Kf-21 is dated with five MC-ICPMS U-Th dates with errors smaller than similar to 1 ka. Kf-21 started forming at similar to 130.2 ka and the main growth phase with relatively constant growth rates lasted from 127.3 ka to 124.4 ka, after which calcite formation ceased. Both delta C-13 and delta O-18 show rapid shifts but also trends, with a range of values within their Holocene counterparts from Korallgrottan. Our results indicate an early onset of the LIG in northern Europe with ice-free conditions at similar to 130 ka. Higher growth rates combined with more negative delta O-18 values between similar to 127.3 and 126.8 ka, interpreted here as warmer and more humid conditions, as well as indications of a millennial-scale cold spell centered at 126.2 ka, resemble findings from speleothem records from other parts of Europe, highlighting that these were regional scale climatic patterns.
8.	Helmens, Karin F., et al. (författare) Ice-free conditions in eastern Fennoscandia during early Marine Isotope Stage 3 : lacustrine records 2010 Ingår i: Boreas. - : Wiley. - 0300-9483 .- 1502-3885. ; 39:2, s. 399-409 Tidskriftsartikel (refereegranskat)abstract The traditional notion that Fennoscandia was glaciated throughout Marine Isotope Stages (MIS) 4-2, from c. 70 kyr BP to the deglaciation 15-10 kyr BP ago, has been challenged during the last decade. Recent studies have shown that climate and environmental settings during MIS 3 were more dynamic than previously assumed, and lacustrine sediment bodies indicate open-water conditions for several sites in eastern Fennoscandia. In this study, three sediment sequences from western, eastern and northeast Finland are compared in detail with respect to their chronology, vegetation reconstruction and climatic inferences. OSL-dating places the sediments in early MIS 3. Pollen evidence suggests the presence of isolated birch trees and open birch forest close to the retreating ice margin, in contrast to vegetation reconstructions from central Europe, which indicate tree-less vegetation. Furthermore, reconstructions of climate using transfer functions have yielded surprising results, indicating present-day summer temperatures in northeast Finland. The combined results suggest ice-free and warm conditions in major parts of eastern Fennoscandia in early MIS 3, possibly during Greenland Interstadial (GIS) 14 around 53 kyr BP ago.
9.	Helmens, Karin F., et al. (författare) Large shifts in vegetation and climate during the early weichselian (mis 5d c) inferred from multi proxy evidence at Sokli (Northern Finland) 2012 Ingår i: Quaternary Science Reviews. - : Elsevier BV. - 0277-3791 .- 1873-457X. ; 41, s. 22-38 Tidskriftsartikel (refereegranskat)abstract For decades, detailed studies on Early Weichselian deposits have been made in central Europe. In contrast, these studies are rare in Fennoscandia in northern Europe. We here integrate an extensive multi-proxy data set obtained on sediments of MIS 5d-c age that form part of a long sediment record preserved at Sokli in northern Finland. We make a detailed interpretation of the vegetation and depositional history for MIS 5d-c using pollen, macrofossils, diatoms and other siliceous microfossils, insect remains, and sediment characteristics, and combine these data with recently published estimates on July temperatures based on chironomids and selected plant indicator species in order to make inferences of paleo-climate regimes. The fossil record obtained on the seven meter thick MIS 5d-c deposit at Sokli is exceptionally rich in species due to the large variety of habitats associated with an overall fluvial depositional environment. A braided river system flanked by steppe-tundra vegetation is inferred for MIS 5d. Mean July temperatures of at least 12-14 degrees C are indicated by chironomids and plant indicator species and are in agreement with the presence of conifers and birch trees as recorded by macrofossils. The reconstructed environmental conditions suggest strong continental climate conditions at Sokli during MIS 5d. The gradual infilling of an oxbow lake and subsequent return to stream channel deposition is traced in great detail in the overlying gyttja and gravelly sediment of MIS 5c age. The terrestrial pollen and plant macrofossil record from the gyttja shows the establishment of birch forest followed by the spread of pine and then spruce. Rich plant indicator species assemblages indicate that the boreal environment at Sokli during MIS 5c experienced July temperatures several degrees higher than the present-day value of 13 degrees C. The high summer temperatures and presence of larch suggest more continental conditions. More open vegetation returned at Sokli during late MIS 5c and was followed by glaciation by the Fennoscandian Ice Sheet (MIS 5b). Despite the major differences in zonal vegetation types during MIS 5d-c, differences in reconstructed July temperatures are minor and suggest that winter temperatures combined with precipitation values mostly determined the stadial-interstadial climate variability. The most compelling conclusion from our study is that forest development during MIS 5c was remarkably similar to that recorded for the Eemian (MIS 5e) and Holocene Interglacials at the high-latitude site Sokli, and also to that inferred from MIS 5c sediments on the northern European mainland. Our results question the definition of MIS 5c in the terrestrial record of Europe as an interstadial interval of the last glacial cycle and suggest inclusion of MIS 5c together with the Eemian (MIS 5e) in an interglacial complex.
10.	Helmens, Karin F., et al. (författare) Major cooling intersecting peak Eemian Interglacial warmth in northern Europe 2015 Ingår i: Quaternary Science Reviews. - : Elsevier BV. - 0277-3791 .- 1873-457X. ; 122, s. 293-299 Tidskriftsartikel (refereegranskat)abstract The degree of climate instability on the continent during the warmer-than-present Eemian Interglacial (around ca. 123 kyr ago) remains unsolved. Recently published high-resolution proxy data from the North Atlantic Ocean suggest that the Eemian was punctuated by abrupt events with reductions in North Atlantic Deep Water formation accompanied by sea-surface temperature cooling. Here we present multiproxy data at an unprecedented resolution that reveals a major cooling event intersecting peak Eemian warmth on the North European continent. Two independent temperature reconstructions based on terrestrial plants and chironomids indicate a summer cooling of the order of 2-4 degrees C. The cooling event started abruptly, had a step-wise recovery, and lasted 500-1000 yr. Our results demonstrate that the common view of relatively stable interglacial climate conditions on the continent should be revised, and that perturbations in the North Atlantic oceanic circulation under warmer-than-present interglacial conditions may also lead to abrupt and dramatic changes on the adjacent continent.
11.	Helmens, Karin F. (författare) The Last Interglacial Glacial cycle (MIS 5-2) re-examined based on long proxy records from central and northern Europe 2014 Ingår i: Quaternary Science Reviews. - : Elsevier BV. - 0277-3791 .- 1873-457X. ; 86, s. 115-143 Tidskriftsartikel (refereegranskat)abstract Current multi-proxy studies on a long sediment sequence preserved at Sokli (N Finland), i.e. in the central area of Fennoscandian glaciations, are drastically changing classic ideas of glaciations, vegetation and climate in northern Europe during the Late Pleistocene. The sediments in the Sokli basin have escaped major glacial erosion due to non-typical bedrock conditions. In this review, the Sokli record is compared in great detail with other long proxy records from central, temperate and northern, boreal Europe. These comprise the classic records of La Grande Pile (E France) and Oerel (N Germany) and more recently obtained records from Horoszki Duke (E Poland) and Lake Yamozero (NW Russia). The focus of the review is on pollen, lithology and macrofossil- and insect-based temperature inferences. The long records are further compared with recent proxy data from nearby terrestrial sites as well as with the rapidly accumulating high-resolution proxy data from the ocean realm. The comparison allows a re-examination of the environmental history and climate evolution of the Last Interglacial Glacial (LI-G) cycle (MIS 5-2). It shows that environmental and climate conditions during MIS 5 (ca 130-70 ka BP) were distinctly different from those during MIS 4-2 (ca 70-15 ka BP). MIS 5 is characterized by three long forested intervals (broadly corresponding to MIS 5e, 5c, 5a), both in temperate and northern boreal Europe. These mild periods were interrupted by two short, relatively cold and dry intervals (MIS 5d and 5b) with mountain-centered glaciation in Fennoscandia. Millennial scale climate events were superimposed upon these longer lasting climate fluctuations. The time interval encompassing MIS 4-2 shows open vegetation. It is characterized by two glacial maxima (MIS 4 and 2) with sub-continental scale glaciation over northern Europe and dry conditions in strongly continental eastern European settings. High amplitude climate oscillations of millennial duration characterized the climate variability of MIS 3. Mild climate conditions in early MIS 3 caused large-scale deglaciation of the Fennoscandian Ice Sheet, and ice-free conditions with Betula-dominated vegetation (including tree birch) persisted over large parts of Fennoscandia, possibly interrupted by glaciation, during major part of MIS 3 till ca 35 ka BP. Overall, MIS 5 was mostly mild with warmest or peak interglacial conditions at the very start during MIS 5e. MIS 4-2 was mostly cold with most extreme or peak glacial conditions in the closing phase during MIS 2. This points to a subdivision of the last climate cycle into an early, overall mild interglacial half and a late, overall cold glacial half, each with duration of ca 50 ka. This review also shows that the climate variability in central and northern Europe during the LI-G cycle was mostly in degrees of continentality with major shifts in winter temperature and precipitation values; summer temperatures, on the other hand, remained largely unchanged. It points to the waxing and waning of sea-ice over the North Atlantic Ocean as a possible characteristic feature of the Late Pleistocene. The present compilation, based on long terrestrial sequences, high-resolution multi-proxy data from the oceans, and quantified paleo-climate data, strongly favors a definition of entire Marine Oxygen Isotope Stage 5 as the Last Interglacial similar as in the original marine stratigraphy and the stratigraphy at La Grande Pile in France. The proxy-based climate data places the start of the Last Glacial at the base of MIS 4 and the northwest European Pleniglacial. It shows that the division between the Eemian (MIS 5e) and the Early Weichselian (MIS 5d-a) is not useful, as not relevant from a climate point of view.
12.	Helmens, Karin F., et al. (författare) Warm summers and rich biotic communities during N-Hemisphere deglaciation 2018 Ingår i: Global and Planetary Change. - : Elsevier BV. - 0921-8181 .- 1872-6364. ; 167, s. 61-73 Tidskriftsartikel (refereegranskat)abstract Detailed studies on fossil remains of plants or animals in glacial lake sediments are rare. As a result, environmental conditions right at the moment of deglaciation of the large N-Hemisphere ice-sheets remain largely unknown. Here we study three deglacial phases of the Fennoscandian Ice Sheet as a unique, repeated element in a long sediment record preserved at Soldl in northern Finland. We summarize extensive multi-proxy data (diatoms, phytoliths, chironomids, pollen, spores, non-pollen palynomorphs, macrofossils, lithology, loss-on-ignition, C/N) obtained on glacial lake sediments dated to the early Holocene (ca. 10 kyr BP), early MIS 3 (ca. 50 kyr BP) and early MIS 5a (ca. 80 kyr BP). In contrast to the common view of an unproductive ice-marginal environment, our study reconstructs rich ecosystems both in the glacial lake and along the shores with forest on recently deglaciated land. Higher than present-day summer temperatures are reconstructed based on a large variety of aquatic taxa. Rich biota developed due to the insolation-induced postglacial warming and high nutrient levels, the latter resulting from erosion of fresh bedrock and sediment, leaching of surface soils, decay of plant material under shallow water conditions, and sudden decreases in lake volume. Aquatic communities responded quickly to deglaciation and warm summers and reflect boreal conditions, in contrast to the terrestrial ecosystem which responded with some delay probably due to time required for slow soil formation processes. Birch forest is reconstructed upon deglaciation of the large LGM ice-sheet and shrub tundra following the probably faster melting smaller MIS 4 and MIS 5b ice-sheets. Our study shows that glacial lake sediments can provide valuable palaeo-environmental data, that aquatic biota and terrestrial vegetation rapidly accommodated to new environmental conditions during deglaciation, and that glacial lake ecosystems, and the carbon stored in their sediments, should be included in earth system modeling.
13.	Hessler, Ines, et al. (författare) Millennial-scale changes in vegetation records from tropical Africa and South America during the last glacial 2010 Ingår i: Quaternary Science Reviews. - : Elsevier BV. - 0277-3791 .- 1873-457X. ; 29:21-22, s. 2882-2899 Tidskriftsartikel (refereegranskat)abstract To reconstruct the response of vegetation to abrupt climate changes during the last glacial we have compiled pollen records from the circum-Atlantic tropics between 23 degrees N and 23 degrees S from both marine and terrestrial sediment cores. Pollen data were grouped into mega-biomes to facilitate the comparison between the different records. Most tropical African records do not appear to register Dansgaard-Oeschger (D-O) variability, although there are vegetation changes during Heinrich Stadials (HS). There is a stronger signal of D-O and HS variability in the South American records. Records close to the modern northern and southern limits of the Intertropical Convergence Zone (ITCZ) show opposite trends in vegetation development during HS and D-O cycles. The pollen data from tropical South America corroborate the hypothesis of a southward shift in the migration pattern of the ITCZ and a reduction in the Atlantic Meridional Overturning Circulation (AMOC) during HS.
14.	Plikk, Anna, 1982-, et al. (författare) Chironomid-based temperature reconstruction for the Eemian Interglacial (MIS 5e) at Sokli, northeast Finland 2019 Ingår i: Journal of Paleolimnology. - : Springer Science and Business Media LLC. - 0921-2728 .- 1573-0417. ; 61:3, s. 355-371 Tidskriftsartikel (refereegranskat)abstract The Last Interglacial (Eemian, MIS 5e) can be considered a test-bed for climate dynamics under a warmer-than-present climate. In this study we present a chironomid record from the high latitude Sokli site (N Finland), where a long continuous sediment sequence from the Last Interglacial has been preserved from glacial erosion. The chironomid-analysis shows a diverse fauna, with dominance of warm-water indicators and shifts in assemblage composition that can be attributed to temperature, lake depth, productivity and habitat availability. Quantitative mean July paleotemperature estimates based on the chironomid data indicate overall mean July temperatures up to 1°C warmer than present. Two cooling events can be discerned, the Tunturi event, dated to about 127 ka BP, in the lower part of the sequence, and the Värriö event, dated to about 119 ka BP, associated with the beginning of a cooling trend in the upper part of the record. Warm conditions already at the onset of the interglacial contrast with a recent chironomid-based Last Interglacial temperature reconstruction from Denmark which suggests a late onset of Eemian warming. The relatively small increase in inferred temperatures compared to present day temperatures differs from other high latitude Eemian sites, and likely reflects the influence of the Atlantic Meridional Overturning Circulation in maintaining already elevated temperatures in Fennoscandia during interglacials.
15.	Plikk, Anna, et al. (författare) Development of an Eemian (MIS 5e) Interglacial palaeolake at Sokli (N Finland) inferred using multiple proxies 2016 Ingår i: Palaeogeography, Palaeoclimatology, Palaeoecology. - : Elsevier BV. - 0031-0182 .- 1872-616X. ; 463, s. 11-26 Tidskriftsartikel (refereegranskat)abstract A 12 m long lacustrine record from Sokli, N Finland, was analyzed for diatoms, non-pollen palynomorphs, macrofossils, pollen and geochemistry in order to reconstruct the development of a high-latitude Eemian lake and investigate the influence of climatic and environmental changes on the lake ecosystem. Based on this multi-proxy dataset we distinguished five major lake phases in the lake's evolution. An initial minerogenic, glacio-lacustrine phase was followed by an organic-rich early Eemian lake phase characterized by anoxic bottom waters, high seasonality and rising nutrient levels. A long open water season, pronounced summer stratification and high productivity characterized the following early mid-Eemian lake phase, corresponding to the Eemian thermal maximum. During the late mid-Eemian lake phase decreasing water depths due to infilling and extensive mixing of the water column resulted in less stable summer stratification and decreased anoxia. The late-Eemian lake phase was characterized by shallow and dynamic conditions and a cooling climate. Superimposed on these general trends are two events characterized by colder and more arid conditions, that possibly match cold and arid events registered in palaeolimnological records on the European continent. In general, the multi-proxy record reflects a nutrient rich lake, where changes in mixing regime associated with climatic forcing and lake level changes asserted a major impact on the aquatic assemblages. The changes in the aquatic assemblages reflect the major patterns of climate change that took place during the Eemian in northern Europe; i.e. a rapid warming and high seasonality during the early Eemian, decreased seasonality during the mid Eemian and a cooling late Eemian with increased seasonality. The high latitude Sokli Eemian palaeolake record lengthens the latitudinal extent of Eemian terrestrial records across Europe, adding to the understanding of climatic gradients and drivers over Europe.
16.	Plikk, Anna, 1982-, et al. (författare) Diatom assemblages from an Eemian palaeolake in northern Europe with morphological observations of rare Aulacoseira sp. resting spores Annan publikation (övrigt vetenskapligt/konstnärligt)abstract An overview of the diatom flora of the last interglacial (Eemian) lacustrine gyttja deposit at Sokli, northeastern Finland is presented together with detailed descriptions including photomicrographs of resting spores similar to the Aulacoseira islandica-skvortzowii group, not found in recent material from Europe. Comparisons are made with literature data on similar taxa. The morphology of the resting spores (e.g. the number of striae and areolae in 10 µm) attributes them to the A. islandica-skvortzowii group but the lack of vegetative cells in the Sokli record makes declaration of true synonymy difficult. The presence of spores similar/related to the A. islandica-skvortzowii group in the Sokli Eemian palaeolake may reflect a more widespread distribution of this group in the past.
17.	Plikk, Anna, 1982-, et al. (författare) Diatom assemblages from an Eemian palaeolake in Northern Europe with morphological observations of rare Aulacoseira sp. resting spores 2021 Ingår i: Diatom Research. - 0269-249X .- 2159-8347. ; 36:4, s. 313-321 Tidskriftsartikel (refereegranskat)abstract An overview of the diatom flora of the last interglacial (Eemian) lacustrine gyttja deposit at Sokli, northeastern Finland is presented together with descriptions including photomicrographs of resting spores similar to the Aulacoseira islandica-skvortzowii group, not found in recent material from Europe. Comparisons are made with literature data on similar taxa. The morphology of the resting spores (e.g., the number of striae and areolae in 10 mu m) links them to the A. islandica-skvortzowii group, but the lack of vegetative cells in the Sokli record makes the determination of true synonymy difficult. The presence of spores similar/related to the A. islandica-skvortzowii group in the Sokli Eemian palaeolake may reflect a more widespread distribution of this group in the past.
18.	Plikk, Anna, 1982- (författare) The Eemian Interglacial at Sokli, northern Finland : A multi-proxy environmental and climatic reconstruction based on a 9 m long lacustrine sediment sequence 2018 Doktorsavhandling (övrigt vetenskapligt/konstnärligt)abstract Understanding the fundamentals of the Earth’s climate system during the Quaternary Period (the past ca 2.6 million years) is of imminent interest in light of the reported recent climatic changes. The Eemian Interglacial (ca 129,000-116,000 years ago) is the last period before the present interglacial (the Holocene, the last ca 11,700 years) when temperatures were as warm as or warmer than today and may serve as an analogue to future climatic warming. The Eemian is a relatively well-studied period, but many aspects of the interglacial climatic dynamics still remain unresolved. One reason for this is the chronological uncertainties of many records from this time period; another is the relatively uneven distribution of palaeoarchives. Especially from high latitude areas, terrestrial records from the Eemian are few due to the impact of glacial erosion during the Weichselian Glacial (ca 116,000-11,700 years ago).This thesis is based on data from Sokli, N Finland, where cold-based conditions of the Weichselian ice sheets and non-typical bedrock conditions have allowed preservation of an unusually thick Eemian lacustrine deposit in stratigraphic sequence with overlying Weichselian and Holocene deposits. The main purpose of this thesis is to contribute to the knowledge of climate variability in Fennoscandia and the North Atlantic region during the Eemian, and it involves a qualitative multi-proxy reconstruction of the Eemian lake and its development as well as a quantitative reconstruction of the regional Eemian July temperature evolution using subfossil chironomids. Attention is paid also to the identification and characterization of short-term climatic events in the Sokli record.The main contribution of this study is to increase the spatial coverage of Eemian palaeoclimatological data. Specific results include the description of the lake development, identifying phases where non-climatic variables likely affect the quantitative chironomid-based temperature reconstruction, as well as qualitative and quantitative reconstructions of climatic features such as changes in seasonality and temperature. Five major lake phases are distinguished. A glaciolacustrine phase is followed by an early Eemian lake phase characterized by short mixing periods and rising nutrient levels. The following early mid-Eemian lake phase is characterized by a prolonged open water season and high productivity and possibly corresponds to the Eemian climatic optimum. The late mid-Eemian lake phase is characterized by infilling and eutrophic conditions while the late Eemian lake phase reflects shallow, dynamic and cooling conditions.The qualitative study suggests a shift from a higher seasonality in the early Eemian towards more oceanic conditions during the mid-Eemian, while chironomid-inferred July temperatures indicate that near present-day temperatures prevailed at Sokli already from the beginning of the interglacial and that optimum temperatures were only moderately (ca + 1°C) higher than at present. These results suggest that the climatic development at Sokli followed that of central Europe during the Eemian. Two events characterized by colder and more arid conditions are identified, and can be tentatively correlated to meltwater events registered in marine records from the Nordic seas and subpolar north Atlantic, possibly reflecting the effect that meltwater pulses and related disruptions in the thermohaline circulation had on adjacent continental areas.
19.	Salonen, J. Sakari, et al. (författare) Abrupt high-latitude climate events and decoupled seasonal trends during the Eemian 2018 Ingår i: Nature Communications. - : Springer Science and Business Media LLC. - 2041-1723. ; 9 Tidskriftsartikel (refereegranskat)abstract The Eemian (the Last Interglacial; ca. 129-116 thousand years ago) presents a testbed for assessing environmental responses and climate feedbacks under warmer-than-present boundary conditions. However, climate syntheses for the Eemian remain hampered by lack of data from the high-latitude land areas, masking the climate response and feedbacks in the Arctic. Here we present a high-resolution (sub-centennial) record of Eemian palaeoclimate from northern Finland, with multi-model reconstructions for July and January air temperature. In contrast with the mid-latitudes of Europe, our data show decoupled seasonal trends with falling July and rising January temperatures over the Eemian, due to orbital and oceanic forcings. This leads to an oceanic Late-Eemian climate, consistent with an earlier hypothesis of glacial inception in Europe. The interglacial is further intersected by two strong cooling and drying events. These abrupt events parallel shifts in marine proxy data, linked to disturbances in the North Atlantic oceanic circulation regime.
20.	Salonen, J. Sakari, et al. (författare) Pollen-based palaeoclimate reconstructions over long glacialinterglacial timescales : methodological tests based on the Holocene and MIS 5dc deposits at Sokli, northern Finland 2013 Ingår i: Journal of Quaternary Science. - : Wiley. - 0267-8179 .- 1099-1417. ; 28:3, s. 271-282 Tidskriftsartikel (refereegranskat)abstract Palaeoclimatic reconstructions over long glacialinterglacial timescales present major methodological challenges, as calibration methods based on modern climate and vegetation patterns may yield biased reconstructions from non-analogue palaeoclimates, such as severely continental glacial periods. In this work, we present pollen-based summer temperature reconstructions based on Holocene and early Weichselian (Marine Isotope Stages (MIS) 5dc) deposits at Sokli, northern Finland. We attempt to improve the robustness of reconstructions of continental palaeoclimates by using two calibration sets, with one located in the region surrounding the fossil site and another in a more continental region in northwestern Russia. We assess the reliability of the reconstructions by estimating the compositional fit between fossil and calibration samples and by comparison with independent, plant macrofossil-based reconstructions. We find the fossil samples to fit the extra-regional, high-continentality calibration set better during the early Holocene and MIS 5dc. Especially in MIS 5dc, however, even the high-continentality calibration set fits the fossil samples poorly compared to the fit achieved in the mid and late Holocene. Thus our results highlight the problem of finding applicable calibration data for pre-Holocene periods. We finally discuss the relative strengths and vulnerabilities of different calibration methods in reconstruction of non-analogue palaeoclimates.
21.	Salonen, J. Sakari, et al. (författare) Uncovering Holocene climate fluctuations and ancient conifer populations : Insights from a high-resolution multi-proxy record from Northern Finland 2024 Ingår i: Global and Planetary Change. - 0921-8181 .- 1872-6364. ; 237 Tidskriftsartikel (refereegranskat)abstract A series of abrupt climate events linked to circum-North Atlantic meltwater forcing have been recognised in Holocene paleoclimate data. To address the paucity of proxy records able to characterise robustly the regional impacts of these events, we retrieved a sub-centennial resolution, well-dated core sequence from Lake Kuutsjarvi, northeast Finland. By analysing a range of paleo-environmental proxies (pollen, plant sedimentary ancient DNA, plant macrofossils, conifer stomata, and non-pollen palynomorphs), and supported with proxy-based paleotemperature and moisture reconstructions, we unravel a well-defined sequence of vegetation and climate dynamics over the early-to-middle Holocene. The birch-dominated pioneer vegetation stage was intersected by two transient tree-cover decrease events at 10.4 and 10.1 thousand years ago (ka), likely representing a two-pronged signal of the 10.3 ka climate event. Our data also show a clear signal of the 8.2 ka climate event, previously not well recorded in the European Arctic, with a collapse of the pine-birch forest and replacement by juniper developing in tight synchrony with Greenland isotopic proxies over 8.4-8.0 ka. Supported by climate modelling, severe winter cooling rather than summer might have been driving vegetation disruptions in the early Holocene. The Kuutsjarvi data indicate an early arrival of Norway spruce (Picea abies) by 9.2 ka (pollen, DNA, and stoma finds), as well as the first evidence for Holocene presence of larch (Larix) in Finland, with pollen finds dating to 9.6-5.9 ka.
22.	Shala, Shyhrete, et al. (författare) Assessment of quantitative Holocene temperature reconstructions using multiple proxies from the Lake Loitsana sediment record Annan publikation (övrigt vetenskapligt/konstnärligt)
23.	Shala, Shyhrete, et al. (författare) Comparison of quantitative Holocene temperature reconstructions using multiple proxies from a northern boreal lake 2017 Ingår i: The Holocene. - : SAGE Publications. - 0959-6836 .- 1477-0911. ; 27:11, s. 1745-1755 Tidskriftsartikel (refereegranskat)abstract Four biotic proxies (plant macrofossils, pollen, chironomids and diatoms) are employed to quantitatively reconstruct variations in mean July air temperatures (T-jul) at Lake Loitsana (northern Finland) during the Holocene. The aim is to evaluate the robustness and biases in these temperature reconstructions and to compare the timing of highest T-jul in the individual reconstructions. The reconstructed T-jul values are evaluated in relation to local-scale/site-specific processes associated with the Holocene lake development at Loitsana as these factors have been shown to significantly influence the fossil assemblages found in the Lake Loitsana sediments. While pollen-based temperatures follow the classical trend of gradually increasing early-Holocene T-jul with a mid-Holocene maximum, the aquatic/wetland assemblages reconstruct higher-than-present T-jul already during the early Holocene, that is, at the peak of summer insolation. The relatively low early-Holocene July temperatures recorded by the pollen are the result of site-specific factors possibly combined with a delayed response of the terrestrial ecosystem compared with the aquatic ecosystem. Our study shows that all reconstructions are influenced at least to some extent by local factors. This finding stresses the need to evaluate quantitatively reconstructed climate values against local lake development and highlights the benefit of using multi-proxy data in Holocene climate reconstructions.
24.	Shala, Shyhrete, et al. (författare) Evaluating environmental drivers of Holocene changes in water chemistry and aquatic biota composition at Lake Loitsana, NE Finland 2014 Ingår i: Journal of Paleolimnology. - : Springer Science and Business Media LLC. - 0921-2728 .- 1573-0417. ; 52:4, s. 311-329 Tidskriftsartikel (refereegranskat)abstract This study presents a detailed analysis of geochemical and biotic proxies in a lake sediment profile to assess the effects of local and regional environmental drivers on the Holocene development of Lake Loitsana, situated in the northern boreal forest of NE Finland. Multi-proxy studies, in particular those that include a detailed plant macrofossil record, from the part of the northern boreal zone of Fennoscandia which has not been affected by treeline fluctuations, are scarce and few of these records date back to the earliest part of the Holocene. A 9-m sediment sequence of gyttja overlying silts representing the last c. 10,700 cal year, allowed for a high-resolution study with emphasis on the early to mid-Holocene lake history. The lacustrine sediments were studied using lithology, loss-on-ignition and C/N ratios, micro- and macro-fossils of aquatic and wetland taxa, diatoms, chironomids and accelerator mass spectrometry C-14 dating on terrestrial plant macrofossils. Our study shows that the local development at Loitsana was complex and included a distinct glacial lake phase and subsequent drainage, a history of fluvial input affected by nearby wetland expansion, and lake infilling in an eventual esker-fed shallow lake. Enhanced trophic conditions, due to morphometric eutrophication, are recorded as Glacial Lake Sokli drained and open water conditions became restricted to a relatively small Lake Loitsana depression. pH appears to have been stable throughout the Holocene with a well-buffered lake due to the local carbonatite bedrock (Sokli Carbonatite Massif). The fossil assemblage changes are best explained by a complex mixture of drivers, including water-body conditions (i.e. depth, turbidity and turbulence), rate of sediment input, and the general infilling of the lake, highlighting the need to carefully evaluate the possible influence of such local factors as palaeoenvironmental conditions are reconstructed based on aquatic proxies.
25.	Shala, Shyhrete, et al. (författare) Palaeoenvironmental record of glacial lake evolution during the early Holocene at Sokli, NE Finland 2014 Ingår i: Boreas. - : Wiley. - 0300-9483 .- 1502-3885. ; 43:2, s. 362-376 Tidskriftsartikel (refereegranskat)abstract The development of a glacial lake impounded along the retreating, northeastern ice margin of the Fennoscandian Ice Sheet during the last deglaciation and environmental conditions directly following the early Holocene deglaciation have been studied in NE Finland. This so-called Sokli Ice Lake has been reconstructed previously using topographic and geomorphologic evidence. In this paper a multiproxy approach is employed to study a 3-m-thick sediment succession consisting of laminated silts grading into gyttja cored in Lake Loitsana, a remnant of the Sokli Ice Lake. Variations in the sediment and siliceous microfossil records indicate distinct changes in water depth and lake size in the Loitsana basin as the Sokli Ice Lake was drained through various spillways opening up along the retreating ice front. Geochemical data (XRF core-scanning) show changes in the influence of regional catchment geochemistry (Precambrian crystalline rocks) in the glacial lake drainage area versus local catchment geochemistry (Sokli Carbonatite Massif) within the Lake Loitsana drainage area during the lake evolution. Principal component analysis on the geochemical data further suggests that grain-size is an additional factor responsible for the variability of the sediment geochemistry record. The trophic state of the lake changed drastically as a result of morphometric eutrophication once the glacial lake developed into Lake Loitsana. The AMS radiocarbon dating on tree birch seeds found in the glaciolacustrine sediment indicates that Lake Loitsana was deglaciated sometime prior to 10 700 cal. a BP showing that tree Betula was present on the deglaciated land surrounding the glacial lake. Although glacial lakes covered large areas of northern Finland during the last deglaciation, only few glaciolacustrine sediment successions have been studied in any detail. Our study shows the potential of these sediments for multiproxy analysis and contributes to the reconstruction of environmental conditions in NE Finland directly following deglaciation in the early Holocene.

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